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The hubs of orb webs, where the spiders lurk, are usually above the center, as the spiders can move downwards faster than upwards. If there is an obvious direction in which the spider can retreat to avoid its own predators, the hub is usually offset towards that direction.
Horizontal orb webs are fairly common, despite being less effective at intercepting and retaining prey and more vulnerable to damage by rain and falling debris.
Various researchers have suggested that horizontal webs offer compensating advantages, such as reduced vulnerability to wind damage; reduced visibility to prey flying upwards, because of the backlighting from the sky; enabling oscillations to catch insects in slow horizontal flight.
However, there is no single explanation for the common use of horizontal orb webs. Spiders often attach highly visible silk bands, called decorations or stabilimenta, to their webs.
Field research suggests that webs with more decorative bands captured more prey per hour. There are several unusual variants of orb web, many of them convergently evolved, including: attachment of lines to the surface of water, possibly to trap insects in or on the surface; webs with twigs through their centers, possibly to hide the spiders from predators; "ladderlike" webs that appear most effective in catching moths.
However, the significance of many variations is unclear. In , Skylab 3 took two orb-web spiders into space to test their web-spinning capabilities in zero gravity.
At first, both produced rather sloppy webs, but they adapted quickly. Members of the family Theridiidae weave irregular, tangled, three-dimensional webs, popularly known as cobwebs.
There seems to be an evolutionary trend towards a reduction in the amount of sticky silk used, leading to its total absence in some species.
The construction of cobwebs is less stereotyped than that of orb-webs, and may take several days. The Linyphiidae generally make horizontal but uneven sheets, with tangles of stopping threads above.
Insects that hit the stopping threads fall onto the sheet or are shaken onto it by the spider, and are held by sticky threads on the sheet until the spider can attack from below.
Although the fossil record of spiders is considered poor,  almost species have been described from fossils. In addition to preserving spiders' anatomy in very fine detail, pieces of amber show spiders mating, killing prey, producing silk and possibly caring for their young.
Hence Attercopus and the similar Permian arachnid Permarachne may not have been true spiders, and probably used silk for lining nests or producing egg cases rather than for building webs.
Several Carboniferous spiders were members of the Mesothelae , a primitive group now represented only by the Liphistiidae.
Some Triassic mygalomorphs appear to be members of the family Hexathelidae , whose modern members include the notorious Sydney funnel-web spider , and their spinnerets appear adapted for building funnel-shaped webs to catch jumping insects.
Araneomorphae account for the great majority of modern spiders, including those that weave the familiar orb-shaped webs. The Jurassic and Cretaceous periods provide a large number of fossil spiders, including representatives of many modern families.
The spiders Araneae are monophyletic i. Arachnids lack some features of other chelicerates, including backward-pointing mouths and gnathobases "jaw bases" at the bases of their legs;  both of these features are part of the ancestral arthropod feeding system.
Pycnogonida sea spiders. Xiphosura horseshoe crabs. Araneae spiders. Pedipalpi whip scorpions , etc. The cladogram shows the relation among spider suborders and families: .
Spiders are divided into two suborders, Mesothelae and Opisthothelae , of which the latter contains two infraorders, Mygalomorphae and Araneomorphae.
Over 48, living species of spiders order Araneae have been identified and as of grouped into families and about 4, genera by arachnologists.
The only living members of the primitive Mesothelae are the family Liphistiidae , found only in Southeast Asia , China , and Japan.
Members of the genus Liphistius run silk " tripwires " outwards from their tunnels to help them detect approaching prey, while those of the genus Heptathela do not and instead rely on their built-in vibration sensors.
The extinct families Arthrolycosidae , found in Carboniferous and Permian rocks, and Arthromygalidae , so far found only in Carboniferous rocks, have been classified as members of the Mesothelae.
However, mygalomorphs cannot produce the pirifom silk that the Araneomorphae use as an instant adhesive to glue silk to surfaces or to other strands of silk, and this makes web construction more difficult for mygalomorphs.
Since mygalomorphs rarely "balloon" by using air currents for transport, their populations often form clumps. Although spiders are widely feared, only a few species are dangerous to people.
Their venom, although they rarely inject much, has resulted in 13 attributed human deaths over 50 years.
There were about reliably reported deaths from spider bites in the 20th century,  compared to about 1, from jellyfish stings.
Even when verification had occurred, details of the treatment and its effects were often lacking. Spider venoms may be a less polluting alternative to conventional pesticides , as they are deadly to insects but the great majority are harmless to vertebrates.
Australian funnel web spiders are a promising source, as most of the world's insect pests have had no opportunity to develop any immunity to their venom, and funnel web spiders thrive in captivity and are easy to "milk".
It may be possible to target specific pests by engineering genes for the production of spider toxins into viruses that infect species such as cotton bollworms.
The Ch'ol Maya use a beverage created from the tarantula species Brachypelma vagans for the treatment of a condition they term 'tarantula wind', the symptoms of which include chest pain, asthma and coughing.
Possible medical uses for spider venoms are being investigated, for the treatment of cardiac arrhythmia ,  Alzheimer's disease ,  strokes ,  and erectile dysfunction.
Spiders can also be used as food. Arachnophobia is a specific phobia —it is the abnormal fear of spiders or anything reminiscent of spiders, such as webs or spiderlike shapes.
Spiders have been the focus of stories and mythologies of various cultures for centuries. In some cultures, spiders have symbolized patience due to their hunting technique of setting webs and waiting for prey, as well as mischief and malice due to their venomous bites.
Web-spinning also caused the association of the spider with creation myths, as they seem to have the ability to produce their own worlds. The Moche people of ancient Peru worshipped nature.
From Wikipedia, the free encyclopedia. For other uses, see Spider disambiguation. Order of arachnids. Main article: Spider anatomy.
Main article: Spider silk. Further information: Spider cannibalism. Play media. Spider fertilization systems. Schematic diagrams showing sperm entering and being stored in the spermathecae; eggs leaving the ovaries and being fertilized; and finally a fertilized egg leaving the female's body.
Main article: Spider web. Main article: Social spider. Main article: Spider evolution. Main article: Spider taxonomy. Main article: Mesothelae.
Main article: Mygalomorphae. Main article: Araneomorphae. Main article: Spider bite. Main article: Arachnophobia. Main article: Cultural depictions of spiders.
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